********************************************************************************
MEME - Motif discovery tool
********************************************************************************
MEME version 5.5.3 (Release date: Sat Jun 10 16:19:38 2023 -0700)

For further information on how to interpret these results please access https://meme-suite.org/meme.
To get a copy of the MEME Suite software please access https://meme-suite.org.

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********************************************************************************
REFERENCE
********************************************************************************
If you use this program in your research, please cite:

Timothy L. Bailey and Charles Elkan,
"Fitting a mixture model by expectation maximization to
discover motifs in biopolymers",
Proceedings of the Second International Conference on Intelligent Systems
for Molecular Biology, pp. 28-36, AAAI Press, Menlo Park, California, 1994.
********************************************************************************


********************************************************************************
TRAINING SET
********************************************************************************
PRIMARY SEQUENCES= common/lipocalin.s
CONTROL SEQUENCES= --none--
ALPHABET= ACDEFGHIKLMNPQRSTVWY
Sequence name            Weight Length  Sequence name            Weight Length  
-------------            ------ ------  -------------            ------ ------  
ICYA_MANSE               1.0000    189  LACB_BOVIN               1.0000    178  
BBP_PIEBR                1.0000    173  RETB_BOVIN               1.0000    183  
MUP2_MOUSE               1.0000    180  
********************************************************************************

********************************************************************************
COMMAND LINE SUMMARY
********************************************************************************
This information can also be useful in the event you wish to report a
problem with the MEME software.

command: meme -p 4 common/lipocalin.s -oc results/meme17 -mod zoops -protein -nmotifs 2 -objfun classic -minw 8 -nostatus 

model:  mod=         zoops    nmotifs=         2    evt=           inf
objective function:           em=       E-value of product of p-values
                              starts=   E-value of product of p-values
width:  minw=            8    maxw=           50
nsites: minsites=        2    maxsites=        5    wnsites=       0.8
theta:  spmap=         pam    spfuzz=        120
em:     prior=       megap    b=            4515    maxiter=        50
        distance=    1e-05
trim:   wg=             11    ws=              1    endgaps=       yes
data:   n=             903    N=               5
sample: seed=            0    hsfrac=          0
        searchsize=    903    norand=         no    csites=       1000
Dirichlet mixture priors file: prior30.plib
Letter frequencies in dataset:
A 0.072 C 0.0288 D 0.0687 E 0.0775 F 0.0432 G 0.0576 H 0.0255 I 0.0476 K 0.0864 
L 0.0875 M 0.0177 N 0.0532 P 0.0321 Q 0.0288 R 0.031 S 0.0587 T 0.0476 V 0.0698 
W 0.0166 Y 0.0498 
Background letter frequencies (from file dataset with add-one prior applied):
A 0.0715 C 0.0293 D 0.0683 E 0.0769 F 0.0433 G 0.0574 H 0.026 I 0.0477 K 0.0856 
L 0.0867 M 0.0184 N 0.0531 P 0.0325 Q 0.0293 R 0.0314 S 0.0585 T 0.0477 V 0.0693 
W 0.0173 Y 0.0498 
Background model order: 0
********************************************************************************


********************************************************************************
MOTIF GACPEVKPVENFDISKFAGTWYEIAK MEME-1	width =  26  sites =   5  llr = 244  E-value = 5.0e-006
********************************************************************************
--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 Description
--------------------------------------------------------------------------------
Simplified        A  :6::2:::::::::22:6:2::2:8:
pos.-specific     C  2:4:::::::::::::::::::::::
probability       D  ::::2::::22:8:::::::::::::
matrix            E  ::222::::2::::2::::2::4:::
                  F  :::::2:::::8::::4:::::::::
                  G  4:::::::2:2:::::::a:::::::
                  H  2::::::::::::::::2:::4::::
                  I  :::2:::::::::2::2::::::42:
                  K  ::::::4:22:::2:4:::2:::::6
                  L  ::2::::::::2:2:::::::::2:2
                  M  ::::::::2::::::::::::::2:2
                  N  :::::::::26:2::2:2::::::::
                  P  :::4:::4::::::::::::::::::
                  Q  2:::::2:::::::2:::::::::::
                  R  :2::::2::2:::::2::::::::::
                  S  :::2222:::::::4:::::::2:::
                  T  :::::2:4:::::::::::4::2:::
                  V  ::2:24:24::::2::2::::::2::
                  W  :::::::::::::2::::::a2::::
                  Y  :2::::::::::::::2::::4::::

         bits    5.9                     *     
                 5.3                     *     
                 4.7                     *     
                 4.1                   * *     
Relative         3.5            **     * **    
Entropy          2.9 **     *  ***    ** **  * 
(70.4 bits)      2.3 **** **** *** ***** ** ***
                 1.8 **************************
                 1.2 **************************
                 0.6 **************************
                 0.0 --------------------------

Multilevel           GACPAVKPVDNFDISKFAGTWHEIAK
consensus            CREEDFQTGEDLNKAAIH A YALIL
sequence             HYLIESRVKKG  LENVN E WSM M
                     Q VSSTS MN   VQRY  K  TV  
                         V    R   W            
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 sites sorted by position p-value
--------------------------------------------------------------------------------
Sequence name             Start   P-value                       Site         
-------------             ----- ---------            --------------------------
BBP_PIEBR                     6  1.76e-24      NVYHD GACPEVKPVDNFDWSNYHGKWWEVAK YPNSVEKYGK
ICYA_MANSE                    7  2.29e-24     GDIFYP GYCPDVKPVNDFDLSAFAGAWHEIAK LPLENENQGK
RETB_BOVIN                    4  1.97e-19        ERD CRVSSFRVKENFDKARFAGTWYAMAK KDPEGLFLQD
LACB_BOVIN                   15  2.92e-18 LLALALTCGA QALIVTQTMKGLDIQKVAGTWYSLAM AASDISLLDA
MUP2_MOUSE                   17  6.60e-17 LCLGLTLVCV HAEEASSTGRNFNVEKINGEWHTIIL ASDKREKIED
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 block diagrams
--------------------------------------------------------------------------------
SEQUENCE NAME            POSITION P-VALUE  MOTIF DIAGRAM
-------------            ----------------  -------------
BBP_PIEBR                         1.8e-24  5_[1]_142
ICYA_MANSE                        2.3e-24  6_[1]_157
RETB_BOVIN                          2e-19  3_[1]_154
LACB_BOVIN                        2.9e-18  14_[1]_138
MUP2_MOUSE                        6.6e-17  16_[1]_138
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 in BLOCKS format
--------------------------------------------------------------------------------
BL   MOTIF GACPEVKPVENFDISKFAGTWYEIAK width=26 seqs=5
BBP_PIEBR                (    6) GACPEVKPVDNFDWSNYHGKWWEVAK  1 
ICYA_MANSE               (    7) GYCPDVKPVNDFDLSAFAGAWHEIAK  1 
RETB_BOVIN               (    4) CRVSSFRVKENFDKARFAGTWYAMAK  1 
LACB_BOVIN               (   15) QALIVTQTMKGLDIQKVAGTWYSLAM  1 
MUP2_MOUSE               (   17) HAEEASSTGRNFNVEKINGEWHTIIL  1 
//

--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 position-specific scoring matrix
--------------------------------------------------------------------------------
log-odds matrix: alength= 20 w= 26 n= 778 bayes= 8.22029 E= 5.0e-006 
   -99    144   -149   -133   -258    269    220   -210   -126   -235   -128    -87   -179    195    -34    -79   -105   -210   -298   -234 
   289   -172   -248   -188   -180   -145   -158   -164   -148   -197    -97   -189   -221    -78    150    -45   -105   -120   -261     74 
   -84    312   -314     44   -111   -230   -180     36   -244     92     33   -225   -204   -122   -125   -133    -66    134   -214   -197 
   -92   -302   -187     36   -266   -208   -155     70   -160   -213   -130   -170    388    -48    -75     72   -103   -192   -332   -302 
   115   -282     98    109   -244   -150    -77   -177    -69   -197    -86    -77   -126     44      3    111    -51     64   -285   -236 
   -68   -167   -316   -260    114   -226   -183     30   -258    -72     12   -219   -200   -135   -133     94    163    231   -223   -188 
  -144   -348   -210   -121   -328   -216    -91   -243    200   -239   -133   -129   -194    225    247    114   -111   -243   -308   -284 
   -92   -241   -296   -256   -247   -224   -228   -114   -258   -190   -105   -229    361   -140   -151    -72    242     70   -329   -324 
   -63   -187   -232   -163   -140     75   -134     22     37    -77    241   -166   -172    -50    -40   -100    -53    221   -226   -208 
   -71   -310    103    103   -254   -142    -60   -205     81   -207    -94    124   -126     52    201    -34    -50   -195   -285   -234 
  -242   -375     91   -196   -382     91    -98   -352   -244   -408   -311    349   -242   -122   -182    -80   -157   -389   -396   -351 
  -318   -290   -467   -456    425   -424   -363   -172   -484    -47   -115   -430   -331   -374   -362   -296   -352   -245   -202   -103 
  -301   -418    363   -117   -408   -310   -244   -367   -403   -415   -323    -21   -370   -260   -284   -260   -313   -381   -414   -410 
  -107   -180   -350   -287   -106   -249   -200    252     -6     95     46   -249   -222   -152   -148   -153    -77    137    216   -191 
    96   -298    -79    114   -258   -152    -75   -202    -65   -209    -96    -73   -125    216      7    200    -40   -190   -293   -244 
    64   -339   -187   -115   -314   -207    -99   -238    223   -242   -134    100   -185     18    221   -102   -106   -241   -310   -282 
  -181   -226   -378   -340    342   -310   -122     98   -351    -88    -39   -304   -258   -227   -212   -200   -176     42   -146    170 
   273   -215   -127   -140   -272   -123    195   -223   -154   -247   -141    118   -194    -56    -71    -37    -96   -174   -321   -276 
  -199   -393   -301   -366   -446    397   -329   -419   -374   -473   -336   -252   -326   -311   -240   -221   -320   -402   -399   -453 
    89   -295    -91    104   -246   -153    -67   -184     77   -196    -82    -70   -123     52     29    -26    223   -176   -280   -234 
  -388   -388   -438   -441   -134   -384   -353   -366   -445   -245   -239   -397   -380   -302   -275   -375   -361   -359    569   -207 
  -290   -318   -383   -376     73   -361    354   -252   -386   -230   -173   -259   -307   -176   -201   -252   -277   -283    286    292 
   106   -271    -98    173   -260   -150    -94   -198    -87   -214   -101    -85   -134     26    -13    137    156   -184   -300   -255 
  -245   -306   -430   -410   -197   -384   -361    357   -396     52    172   -359   -346   -288   -287   -287   -197    121   -336   -320 
   328   -140   -375   -329   -260   -140   -310     72   -352   -199   -114   -318   -308   -242   -222    -45   -138    -72   -331   -364 
  -192   -313   -316   -228   -235   -288   -185   -128    276     36    217   -224   -256    -77     45   -195   -163   -176   -307   -304 
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 position-specific probability matrix
--------------------------------------------------------------------------------
letter-probability matrix: alength= 20 w= 26 nsites= 5 E= 5.0e-006 
 0.000000  0.200000  0.000000  0.000000  0.000000  0.400000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.600000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.200000 
 0.000000  0.400000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000 
 0.200000  0.000000  0.200000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.200000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.200000  0.400000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.000000  0.000000  0.200000  0.200000  0.200000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.000000  0.400000  0.200000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.200000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000 
 0.000000  0.000000  0.200000  0.200000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.200000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.200000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.600000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.800000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.800000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.200000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.200000  0.000000 
 0.200000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.400000  0.000000  0.000000  0.000000  0.000000 
 0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.200000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.200000 
 0.600000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.200000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.400000 
 0.200000  0.000000  0.000000  0.400000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.200000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.400000  0.000000  0.200000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000 
 0.800000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.600000  0.200000  0.200000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif GACPEVKPVENFDISKFAGTWYEIAK MEME-1 regular expression
--------------------------------------------------------------------------------
[GCHQ][ARY][CELV][PEIS][ADESV][VFST][KQRS][PTV][VGKM][DEKNR][NDG][FL][DN][IKLVW][SAEQ][KANR][FIVY][AHN]G[TAEK]W[HYW][EAST][ILMV][AI][KLM]
--------------------------------------------------------------------------------




Time  0.36 secs.

********************************************************************************


********************************************************************************
MOTIF VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2	width =  42  sites =   2  llr = 218  E-value = 8.6e-004
********************************************************************************
--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 Description
--------------------------------------------------------------------------------
Simplified        A  :::::5::::::5::::::::::::5::::5:::::::::::
pos.-specific     C  :::::::::::::::::a::::::::::::::::::::::::
probability       D  :::::::a::::::::::5:5:a:::::5:::::::::::::
matrix            E  :::::::::::::::::::::5::::::::::::::::::5:
                  F  :5:::::::::::::::::::::::::::5::::::::::::
                  G  ::::::::::::::5::::::::::5:::::::::::::::a
                  H  ::::::::::::::::::::5:::::a::5::::::::::::
                  I  ::::::::::::5a::::::::::::::5:::5:::::::::
                  K  :::::::::a::::::::5::::aa::::::::::5:a::::
                  L  ::::a::::::::::::::::::::::::::::a:::::a::
                  M  ::::::::::::::::::::::::::::::::::::::::::
                  N  ::5:::::5:a:::5:5:::::::::::::::::::::::::
                  P  :5:::::::::::::::::::5::::::::::::::::::::
                  Q  :::::::::::::::::::::::::::5::::::::::::::
                  R  :::::::::::::::::::::::::::::::::::5::::::
                  S  :::::5:::::::::::::::::::::5::::::a:a:::::
                  T  ::::::a:::::::::::::::::::::::::::::::::5:
                  V  a::a::::::::::::::::::::::::::5:5:::::a:::
                  W  ::5::::::::::::::::::::::::::::a::::::::::
                  Y  ::::::::5::a:::a5::a::::::::::::::::::::::

         bits    5.9                                *          
                 5.3                  *        *    *          
                 4.7       *      *   *        *    *          
                 4.1 * **  **  ** * * * *  *   *  * *  * * *  *
Relative         3.5 ***** ****** * *** ****** ** * * ******* *
Entropy          2.9 ******************************************
(157.6 bits)     2.3 ******************************************
                 1.8 ******************************************
                 1.2 ******************************************
                 0.6 ******************************************
                 0.0 ------------------------------------------

Multilevel           VFNVLATDNKNYAIGYNCDYDEDKKAHQDFAWILSKSKVLEG
consensus             PW  S  Y   I N Y K HP   G SIHV V  R    T 
sequence                                                       
                                                               
                                                               
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 sites sorted by position p-value
--------------------------------------------------------------------------------
Sequence name             Start   P-value                               Site                 
-------------             ----- ---------            ------------------------------------------
ICYA_MANSE                  102  7.93e-48 FKFGQRVVNL VPWVLATDYKNYAINYNCDYHPDKKAHSIHAWILSKSKVLEG NTKEVVDNVL
BBP_PIEBR                    98  4.24e-47 LTYGGVTKEN VFNVLSTDNKNYIIGYYCKYDEDKKGHQDFVWVLSRSKVLTG EAKTAVENYL
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 block diagrams
--------------------------------------------------------------------------------
SEQUENCE NAME            POSITION P-VALUE  MOTIF DIAGRAM
-------------            ----------------  -------------
ICYA_MANSE                        7.9e-48  101_[2]_46
BBP_PIEBR                         4.2e-47  97_[2]_34
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 in BLOCKS format
--------------------------------------------------------------------------------
BL   MOTIF VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG width=42 seqs=2
ICYA_MANSE               (  102) VPWVLATDYKNYAINYNCDYHPDKKAHSIHAWILSKSKVLEG  1 
BBP_PIEBR                (   98) VFNVLSTDNKNYIIGYYCKYDEDKKGHQDFVWVLSRSKVLTG  1 
//

--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 position-specific scoring matrix
--------------------------------------------------------------------------------
log-odds matrix: alength= 20 w= 42 n= 698 bayes= 8.44294 E= 8.6e-004 
   -79   -197   -355   -318   -210   -299   -268    107   -341   -110    -67   -322   -234   -238   -197   -229   -101    331   -339   -344 
  -137   -239   -277   -259    326   -239   -146   -122   -264   -107    -86   -234    308   -139   -142   -141   -155   -151    -95    -24 
  -243   -321   -175   -279   -121   -223    -96   -239   -261   -226   -172    228   -262   -131   -126   -166   -184   -265    489   -152 
   -79   -197   -355   -318   -210   -299   -268    107   -341   -110    -67   -322   -234   -238   -197   -229   -101    331   -339   -344 
  -232   -281   -423   -355    -89   -354   -279     14   -351    312     78   -355   -259   -193   -199   -277   -203    -82   -245   -269 
   260   -124   -231   -211   -237    -73   -197   -179   -228   -221   -115   -161   -123   -111   -115    215     18   -110   -289   -286 
  -115   -208   -264   -265   -263   -222   -217   -136   -243   -236   -102   -136   -191   -130   -130     78    380   -138   -309   -322 
  -283   -407    363    -98   -394   -295   -235   -352   -376   -398   -307    -82   -341   -233   -263   -250   -296   -363   -401   -396 
  -147   -254    -80   -171     21   -153     82   -155   -172   -189   -105    285   -177    -52    -64    -70   -102   -191    -74    197 
  -305   -400   -419   -355   -479   -375   -305   -337    336   -408   -297   -310   -331   -224     52   -330   -294   -410   -407   -460 
  -290   -351   -175   -321   -339   -249    -79   -270   -296   -362   -255    396   -272   -163   -203   -117   -181   -336   -332   -353 
  -177   -240   -286   -271    130   -250     12   -148   -272   -147    -90   -220   -212   -153   -133   -164   -184   -183    -24    366 
   186   -147   -282   -245   -131   -169   -213    265   -247    -37      9   -228   -165   -135   -134    -90    -74    105   -242   -226 
  -227   -291   -398   -381   -193   -360   -348    379   -367    -11      8   -337   -320   -272   -264   -272   -181    100   -334   -306 
  -132   -312    -93   -193   -330    316   -121   -289   -216   -339   -227    220   -205   -115   -118    -85   -149   -292   -313   -322 
  -177   -240   -286   -271    130   -250     12   -148   -272   -147    -90   -220   -212   -153   -133   -164   -184   -183    -24    366 
  -147   -254    -80   -171     21   -153     82   -155   -172   -189   -105    285   -177    -52    -64    -70   -102   -191    -74    197 
  -352    503   -549   -513   -466   -488   -464   -376   -574   -454   -338   -497   -462   -440   -393   -414   -349   -449   -559   -570 
  -129   -328    242     -9   -297   -142   -105   -237    165   -262   -158    -42   -149     -4     21    -88   -109   -238   -296   -276 
  -177   -240   -286   -271    130   -250     12   -148   -272   -147    -90   -220   -212   -153   -133   -164   -184   -183    -24    366 
  -183   -327    287    -65   -253   -171    284   -276   -201   -289   -190     -4   -197    -16    -79   -118   -154   -273   -282   -182 
  -101   -319    -72    213   -284   -181   -145   -229   -148   -241   -161   -148    341     12    -70   -102   -119   -215   -319   -301 
  -283   -407    363    -98   -394   -295   -235   -352   -376   -398   -307    -82   -341   -233   -263   -250   -296   -363   -401   -396 
  -305   -400   -419   -355   -479   -375   -305   -337    336   -408   -297   -310   -331   -224     52   -330   -294   -410   -407   -460 
  -305   -400   -419   -355   -479   -375   -305   -337    336   -408   -297   -310   -331   -224     52   -330   -294   -410   -407   -460 
   199   -186   -237   -248   -298    312   -237   -239   -270   -284   -174   -191   -182   -169   -150    -53   -130   -169   -312   -340 
  -251   -299   -223   -251   -169   -257    478   -292   -278   -251   -155    -70   -221     34    -46   -163   -183   -289   -217    -71 
   -58   -241   -120    -57   -236   -135     -5   -183    -92   -192    -82    -53    -91    354     16    179     18   -185   -243   -234 
  -161   -273    270    -82   -205   -214   -177    215   -233   -119    -74    -88   -202   -113   -137   -161   -140     -8   -273   -263 
  -185   -239   -243   -252    281   -254    348   -145   -265   -129    -82   -121   -202    -31    -84   -154   -166   -174    -42    136 
   212   -142   -296   -257   -186   -157   -226     48   -277   -120    -55   -255   -177   -162   -152    -78    -74    246   -277   -283 
  -372   -376   -426   -428   -122   -371   -335   -347   -429   -233   -225   -383   -367   -289   -261   -360   -346   -344    567   -191 
  -159   -242   -393   -370   -194   -350   -324    329   -369    -32     -7   -335   -296   -267   -250   -262   -144    224   -335   -319 
  -232   -281   -423   -355    -89   -354   -279     14   -351    312     78   -355   -259   -193   -199   -277   -203    -82   -245   -269 
   -69   -199   -243   -264   -279   -169   -219   -249   -253   -287   -177   -124   -169   -158   -135    352    119   -256   -326   -308 
  -247   -353   -346   -266   -414   -302   -180   -299    268   -325   -228   -236   -261    -82    333   -237   -222   -342   -332   -379 
   -69   -199   -243   -264   -279   -169   -219   -249   -253   -287   -177   -124   -169   -158   -135    352    119   -256   -326   -308 
  -305   -400   -419   -355   -479   -375   -305   -337    336   -408   -297   -310   -331   -224     52   -330   -294   -410   -407   -460 
   -79   -197   -355   -318   -210   -299   -268    107   -341   -110    -67   -322   -234   -238   -197   -229   -101    331   -339   -344 
  -232   -281   -423   -355    -89   -354   -279     14   -351    312     78   -355   -259   -193   -199   -277   -203    -82   -245   -269 
   -93   -271    -46    252   -263   -166   -113   -176   -114   -220   -118    -96   -123     29    -40    -20    180   -162   -288   -264 
  -158   -352   -261   -321   -404    391   -289   -372   -331   -425   -294   -212   -283   -265   -201   -180   -272   -356   -365   -412 
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 position-specific probability matrix
--------------------------------------------------------------------------------
letter-probability matrix: alength= 20 w= 42 nsites= 2 E= 8.6e-004 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000 
 0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000 
 0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000 
 0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.500000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.500000  0.000000  0.000000  0.000000 
 0.000000  0.000000  0.000000  0.000000  0.000000  1.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000  0.000000 
--------------------------------------------------------------------------------

--------------------------------------------------------------------------------
	Motif VFWVLATDNKNYIIGYNCDYDPDKKGHQDFVWILSKSKVLEG MEME-2 regular expression
--------------------------------------------------------------------------------
V[FP][NW]VL[AS]TD[NY]KNY[AI]I[GN]Y[NY]C[DK]Y[DH][EP]DKK[AG]H[QS][DI][FH][AV]W[IV]LS[KR]SKVL[ET]G
--------------------------------------------------------------------------------




Time  0.61 secs.

********************************************************************************


********************************************************************************
SUMMARY OF MOTIFS
********************************************************************************

--------------------------------------------------------------------------------
	Combined block diagrams: non-overlapping sites with p-value < 0.0001
--------------------------------------------------------------------------------
SEQUENCE NAME            COMBINED P-VALUE  MOTIF DIAGRAM
-------------            ----------------  -------------
ICYA_MANSE                       6.79e-65  6_[1(2.29e-24)]_69_[2(7.93e-48)]_46
LACB_BOVIN                       1.79e-15  14_[1(2.92e-18)]_138
BBP_PIEBR                        2.22e-64  5_[1(1.76e-24)]_66_[2(4.24e-47)]_34
RETB_BOVIN                       1.28e-16  3_[1(1.97e-19)]_154
MUP2_MOUSE                       2.21e-14  16_[1(6.60e-17)]_138
--------------------------------------------------------------------------------

********************************************************************************


********************************************************************************
Stopped because requested number of motifs (2) found.
********************************************************************************

CPU: Timothys-iMac.local

********************************************************************************
